dependent cell that can only grow with a substratum
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Substratum rule is when there are multiple facets of layers for the rule of whatever is being governed or managed.
The substratum is located below the subsoil. It is mainly composed of medium-sized rocks, usually sediments and Aeolian deposits, and light-colored soil.
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"Foundation" is one definition.Agriculturally speaking, it means subsoil.See the related Online Dictionary link listed below for more information:
Rhizoids
Fungi
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They are filamentous thread like structures which grow horizontally on the surface of the substratum
The substratum is the stratum below the stratum being discussed.
root hairs