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What you are referring to is the sinus. The crayfish's gills are used to extract oxygen that is dissolved in water and lead it into the crayfish's open circulatory system. This type of circulatory system doesn't limit the blood to blood vessels; instead the blood collects in the pericardial sinus, the cavity surrounding the dorsal heart. (Porch and Batdorf 477) From there the dorsal heart pumps the blood through short vessels, which empty into internal cavities and bathe the organs. After the blood bathes the organs and drains from these cavities, it flows into the large sternal sinus. The blood then passes through the gills, becomes oxygenated, and returns to the pericardial. The green glands, located near the base of the antennae, filter out waste as the blood circulates. Then the waste is excreted through a pore just in front of the Crayfish's mouth.

Reference:

Bob Jones University-Thomas E. Porch, and Brad R. Batdorf. Biology. Greenville, South Carolina: BJU Press, 2005.

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Q: What is the cinus in the thorax of a crayfish?
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