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There are a number of issues that caused a considerable drop in temperature, meteor strikes in three locations, a global decrease in CO2 levels (this is the current/leading theory). There is some evidence that the meteor strikes probably increased the cooling effect and probably resulted in the mass extinction of equids and the subsequent migration of ruminant animals from Asia to fill the voids.

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Q: What might have caused the extinction or made it worse at the end of the Eocene epoch?
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