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It depends on use.

The Wizard decreed: There will be NO sharing of shoe laces. All those who participate in this dastardly act will be forced to squish noodles through their teeth. Here, "all those" is plural, so the plural form of the verb is used.

The investigator found irrefutable evidence that a large group of people gathered there and shared bellybutton lint. "Who participates in such heinous things?" The investigator wondered, mystified. Here the use of the word "who" is singular, as in "what sort of individual participates". So the singular form of the verb is used.

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Q: Who participate or who participates?
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