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C4 fixation is an elaboration of the more common C3 carbon fixation and is believed to have evolved more recently. and CAM overcome the tendency of the enzyme RuBisCoto wastefully fix oxygen rather than carbon dioxide in what is called photorespiration. This is achieved by using a more efficient enzyme to fix in mesophyll cells and shuttling this fixed carbon via malate or asparate to bundle-sheath cells. In these bundle-sheath cells, RuBisCO is isolated from atmospheric oxygen and saturated with the released by decarboxilation of the malate or oxaloacetate. These additional steps, however, require more energy in the form of ATP. Because of this extra energy requirement, plants are able to more efficiently fix carbon in only certain conditions, with the more common pathway being more efficient in other conditions.

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Q: What is the purpose of the hatch-slack pathway?
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