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He meant that animals have to expend energy in order for them to gain enough food to survive and reproduce. The less efficient their attempts to gather food, the less energy they'll have for raising fertile offspring, the lower the chances that their genes will be represented in subsequent generations. Through reproduction, animals indirectly compete with all other members of their species, who are also trying to gather food, find mates and get their genes represented in the next generation.

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Q: What did Darwin mean when he said that individuals struggle to survive?
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