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When seeds germinate they are under the ground and as such are not able to detect any light source to grow towards however they are able detect the force of gravity just like humans do. No matter which way round you plant a seed the roots always grow down and the shoot grows up, the roots show positive geotropism (this means they grow in the same direction as the pull of gravity i.e. down) and the shoots show negative geotropism which is against gravity i.e. upwards. As soon as the shoot emerges above the soil it will then grow towards a source of light so that it can convert the suns energy into usable energy through photosynthesis. It is in the plants best interest to grow straight upwards so that it get as much light as it can and not be out competed by other plants.

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