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Cornwallis was the eldest son of Charles Cornwallis, 5th Baron Cornwallis (later 1st Earl Cornwallis) (29 March 1700 - 23 June 1762, in the Howells, near Bristol) and was born at Grosvenor Square in London,England, even though his family's estates were in Kent.

The Cornwallis family was established at Brome Hall, near Eye, in Suffolk, in the 14th century, and its members occasionally represented the county in the House of Commons over the next three hundred years. Frederick Cornwallis, created a Baronet in 1627, fought for King Charles I, and followed King Charles II into exile. He was made Baron Cornwallis, of Eye in the County of Suffolk, in 1661, and his descendants by fortunate marriages increased the importance of the family.

He was extremely well-connected. His mother, Elizabeth Townshend (died 1 December 1785), was the daughter of the 2nd Viscount Townshend and a niece of the Prime Minister, Sir Robert Walpole. His father was created Earl Cornwallis, Viscount Cornwallis and Viscount Brome in 1753, at which point he was styled Viscount Brome. His brother was Admiral Sir William Cornwallis. An uncle, Frederick, was Archbishop of Canterbury and another uncle, Edward, was a leading colonist in Canada.

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