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When the transmission is operating at a 1:1 ratio, also known as direct drive. In most overdrive transmissions, there isn't a gear which hits at exactly a 1:1 ratio. But if you have an older vehicle with a direct drive transmission (like most of the older three speed automatics and four speed manuals are), then your highest gear is likely a 1:1 ratio. What that means is that, for each rotation of the transmission's input shaft, there is one rotation of the output shaft, as opposed to overdrive, where the output shaft spins faster than the input shaft.

Of course, when the vehicle is shut off and not in motion, the engine speed and driveshaft speed are matching then, as well.

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