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A vascular bundle is a part of the transport system in vascular plants. The transport itself happens in vascular tissue, which exists in two forms: xylem andphloem. Both these tissues are present in a vascular bundle, which in addition will include supporting and protective tissues. Also, it is a vein in the leaf that contains conducting tissues.The xylem typically lies adaxial with phloem positioned abaxial. In a stem or root this means that the xylem is closer to the centre of the stem or root while the phloem is closer to the exterior. In a leaf, the adaxial surface of the leaf will usually be the upper side, with the abaxial surface the lower side. This is whyaphids are typically found on the underside of a leaf rather than on the top, since the sugars manufactured by the plant are transported by the phloem, which is closer to the lower surface.

The position of vascular bundles relative to each other may vary considerably: see stele.

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