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The ecosystem should have, in general, effective negative feedback relations between it's elements.

For instance, a simplest ecosystem "predator-prey-limit" possess 1 negative feedback relation:

1. The more predators are, the less prey is. The less predators are, the more prey is.

The other relation in this system is positive feedback:

2. The more prey is, the more predators are. The less prey is, the less predators are.

If there would be no negative feedback relation (1), then population of both predators and prey would reach ecosystem limit (alimentary and/or area) and then extinct.

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