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Early Paleozoic is pretty long time, so you have to be more specific about that. Probably one of the most dominant creatures that ever evolved was Anomalocaris (from Cambrian period). Most of the animals from that time couple of inches long, while Anomalocaris was 2 meters long. Also giant scorpions, like world famous Jaekelopterus rhenaniae were also top predators of it's days.

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