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That would be the "guard cells". The stoma is an opening on the underside of many leaves and each stoma is bordered by a pair of guard cells that open and close the pore to allow for "transpiration" (the passage of water in either gaseous or liquid form into or out of the leaf). In hot, dry weather, the guard cells close to prevent water loss. In humid weather, the guard cells allow the pore to open and water can enter.

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