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That depends on their MOS (occupation). By the way, since the Specialist ranks above E4 were eliminated, it's just "SPC" now, not SP4. As far as duties, they may be similar to those of other lower enlisted (Privates and Privates First Class), or they may be put in a junior NCO (Corporal and Sergeant) position either until they get promoted or else the unit receives an NCO to replace them. A Specialist who earned their rank through time in service and might actually make a career in the military will likely serve either in the latter role, or else help reinforce the command of a junior NCO - a Specialist who just enlisted as that rank for reasons of a college degree will essentially be an overpaid private.

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