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Muzzle velocity and muzzle energy. 7mm uses .284 diameter bullets and the .270 uses .277 diameter bullets. The .270 is basically a necked down .30-06 case. 7mm comes in several different flavors. The 7mm Mauser is 6mm shorter than the .270/.30-06 and has less powder capacity. the 7mm Remington express (.280 Rem) has the same length (very close) as the .30-06, but the shoulder is slightly farther forward. This prevents it from being accidentally chambered in a .270 or .30-06 and gives a bit more powder capacity.

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