"Foundation" is one definition.
Agriculturally speaking, it means subsoil.
See the related Online Dictionary link listed below for more information:
dependent cell that can only grow with a substratum
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Substratum rule is when there are multiple facets of layers for the rule of whatever is being governed or managed.
The substratum is located below the subsoil. It is mainly composed of medium-sized rocks, usually sediments and Aeolian deposits, and light-colored soil.
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A substance or layer that underlies something, or on which some process occurs, in particular.The surface or material on or from which an organism lives, grows, or obtains its nourishment.
Rhizoids
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Fungi
They are filamentous thread like structures which grow horizontally on the surface of the substratum
The substratum is the stratum below the stratum being discussed.
It the underlying reason a company exists; the foundational purpose of the company.