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No. As soon as you pick up the skull (be sure to pick it up), it will be recorded as found. For some skulls, there is an achievement associated with finding it. As soon as you see this achievement, the skull is considered found. You might want to finish the level anyways, just for fun, but it is not necessary.

I am 100% sure of this. I have tried it both ways, and both ways I get the skull.

P.S. Try to get the skull on Floodgate. It's great.

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Q: To get a halo 3 skull do you have to play the whole mission?
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