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The question itself is actually pertaining to an age-old psychology question of nature versus nurture.

Nature here is the girl's innate qualities. Their biological and emotional make-up such as hormones. The nurture aspect then is the environment that affects a person. For example, these could be the people that the girl has or anything that is external.

So here, the current person is a muscular girl. To consider how that person became a muscular girl, the most accepted idea is that it is a combination of both nature and nurture.

"because their mothers made them that way" would fall into the category of nurture because it was an external environment that had helped shape the person into a muscular girl. BUT it is also possible that a person can be a muscular girl even if their parents did not made them that way. The girl can just be innately more predisposed to being muscular. She can simply have a high productivity of testosterone which would be a determining factor if she would be a muscular girl. This then would be an example of nature.

So to summarize, how a person currently is (whether they are girlie girls or muscular girls or manly boys or feminine boys)would be dependent on their innate qualities (nature) and the external environment (nurture).

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Q: Were muscular girls once feminine girls because their mothers made them that way first?
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