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Since WW2 is considered to have begun on 1 Sept 1939 with the invasion of Poland, this would be the first combat against German air force. The Germans did sent pilots to fly in the Spanish Civil War but they did not fly under German markings.

Section Lt Hieronim Dudwal scored a kill on an He-111 bomber on 1 Sept 1939 while flying a PZL P.11C. Not sure if that was the first one. (See Link) Also on 1 September 1939, Stanisław Skalski,142nd Fighter Squadron, attacked a German Henschel Hs 126 reconnaissance aircraft, which was eventually shot down by Marian Pisarek, and then landed next to it and helped to bandage a crew. He later became an ace. Polish Aces in 1939

http://aces.safarikovi.org/victories/Poland-ww2-pol.pdf List of Polish Aces who flew for Poland and UK, including those who downed V-1 Buzz bombs:

http://aces.safarikovi.org/victories/lista.bajana.pdf Second Answer First German aircraft to be shot down in England was a He111 on the 3rd Feb 1940 from II Gruppe of the ' Lion Geshwader' (KG26). It was shot down by Hurricanes from Blue Section, 43 Sqn. Their leader being Peter Townsend, the other pilots being Folkes and Sgt Hallows. It crash landed a mile or two from Whitby Town. Oberserver Peter Leushake and flight mechanic Johann Meyer were killed. Pilot Hermann Wilms and gunner Karl Missy survived although Karl Missy lost his legs through his injuries.

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Q: WhAT fighter aircraft shot down the first German aircraft and were?
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