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The Germans, by a very wide margin. 85% of the US war effort was directed against Germany, until Germany was defeated. The Pacific War was fought on a shoestring, mainly by the US Navy, the Navy's Marine Corps, which grew to six infantry divisions, and about fifteen army infantry divisions. Everything else was against the Germans.

The Italians also killed a very small percentage of those lost, and the French an even smaller number. The Vichy French were a German ally and fought the US for three or four days after the landings in North Africa in November 1942. Nobody really had their heart in it though, and soon the French came over to our side.

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Q: What killed most Americans in World War 1?
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