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No. Not usually. Look at your tachometer (sp). A car will usually cruise (cruise control) on flat land at about 1.2 to 1.3K rpm at about 70 mph. I have had cars that cruised in this situation at about 9,000 (9K) at 70 mph on overdrive. That simply means that the car is traveling at the same speed on 1 or overdrive, but the motor is working less in overdrive. I got 38.7 miles a gallon on my 1995 Buick LeSabre 6-cylinder. When you come upon an upgrade (hill), notice that the transmission will kick into a stronger gear to make the climb, the rpm increases to keep up but the speed stays the same (70 mph). When it completes the climb, notice that it goes back to overdrive, the tachometer reduces in rpm, but the speed (70 mph) remains the same.

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