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Star Trek is an extensive franchise which includes five different TV series (one being animated) and a series of movies, so there are a number of different arch enemies who appear at different times. In the original series (1966-1968) the arch enemy were the Klingons, who were largely reconciled with the Federation by the time of the second series, Star Trek The Next Generation (which included a Klingon named Worf as a major character on the new Enterprise). Romulans were also arch enemies in the original series, and in parts of other series. Then you get the Cardassians and, later, the Borg emerged as the arch enemy. There were various others as well, but they were prominent for only short story arcs.

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12y ago
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13y ago

Roddenberry used Romulus from the Roman myth of their founding as a basis for the Romulan society. This connection goes deep into the Romulans' government and social structure, as shown by the Romulan Senate and the Praetor. Remus was also used, for the Remans, a neighboring species, but they are shown to be inferior, much the same way Remus was to Romulus before he was slain.

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11y ago

Klingons, Romulans, Borg, Kazon, Vidians & Cardassians.

You could say Q, but they aren't true villains,

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14y ago

Klingons

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Q: Who are the villains of Star Trek?
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