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You can but the vehicle won't run right. The 4L60E helps the engine run correctly. If you look at the 700R4 it has a kick down cable where as the 4L60E doesn't. It is run by the ECM and inputs from the engine. This Trans is totally run by the ECM, it takes care of the shifting points when driving. So yes you can but it won't run right.

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Q: Can you replace a 4l60e trans with a r700 and make it work?
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