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Typically, just age. Each gear of an automatic transmission is operated by one of a set of clutches or bands. When a band or clutch wears out that gear is through and the transmission will need to be rebuilt. The Valve Body controls pressures to different pistons in the transmission. At specific events the valve body causes pressure to be applied to the 2nd gear clutch piston and if the pressure is not sufficient the clutch may "slip", causing additional and premature wear. Driving a vehicle with insufficient ATF can also cause slipping and wear.

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Q: What causes second gear to go bad in an automatic?
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