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In some cases, nothing. Many older model vehicles used to use them interchangeably. Newer model vehicles, however, generally need different fluids. Probably the largest differences are in the fluids' lubricating capabilities and the amount of air/gases dissolved in them. Modern transmissions need a fluid that excells at lubrication, and heat dispersal, while Power Steering applications need better hydraulic compression (a more compressed oil will have less gases in it and will transfer power better) and some lubrication properties.

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Q: What is the difference between automatic transmission oil and power steering oil?
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