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fitness

 
Dictionary: fit·ness   (fĭt'nĭs) pronunciation
n.
  1. The state or condition of being fit; suitability or appropriateness.
  2. Good health or physical condition, especially as the result of exercise and proper nutrition.
  3. Biology. The extent to which an organism is adapted to or able to produce offspring in a particular environment.

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World of the Body: fitness
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Fitness for what?

When we speak, perhaps with a hint of envy, of a ‘fit’ young man or woman — and even more when we refer, with undisguised admiration, to a ‘fit’ old person — there is little ambiguity as to our meaning: we are referring to fitness to cope with life in general, not only with sport, and certainly not a particular sport. Furthermore the international athlete, in peak of condition, is ‘fit’ for only a limited number of very similar events: the sprinter could not possibly run a marathon, the power lifter could compete with neither kind of runner at their events. The fitness of the racing driver is radically different from that of the dinghy sailor, the gymnast from that of the mountaineer and, perhaps most radically of all, the oarsman from that of the pistol shooter. Furthermore, many highly trained athletes, particularly those conditioned for endurance events, display greater, not less, vulnerability than the average person to many forms of illness.

Clearly then, we must distinguish ‘fitness for life’ from ‘fitness for sport’; and, when considering the latter, must specify which sport.

Fitness for life

This is a condition which we almost all desire, but few of us pursue with vigour. To attain and maintain it requires adequate and balanced nourishment, adequate and varied exercise, adequate but not excessive sleep, avoidance of excess in using social drugs, plentiful stimulation without excessive stress, and psychosocial well-being. The Aristotelian precept, ‘moderation in all things’, remains as good a guide as any to the balances which must be struck. Fitness for work, for leisure and recreational exercise, for family life and parenthood, and even for childbearing itself, and fitness to cope with emergencies — all are optimized in these broad ways. The influences of genetics and of environment are inescapable, so the fitness attained by one person will be very different from that attained by another, but all will approach their individual optima by personal application of the same balanced principles. Even Western and Eastern, secular and religious wisdoms (disregarding the most extreme of the latter) have much more in common than divergence in their guidelines for ‘fitness’, whether or not they would recognize that term; and modern science, while adding a few details on matters like trace nutrients, takes little issue with them about the broader picture.

Endurance fitness

If there is one aspect of specialist, sports-oriented fitness which embodies the greatest part of the lay ideal, it is probably endurance fitness — the ability to continue a demanding physical activity many times longer than the untrained person can. Whether the challenge is a London- Brighton cycle race, an ascent of the Matterhorn, or a Channel swim, the fundamentals of this category of fitness are the same. Each of these activities is trained for in essentially the same way — namely, by covering large mileages several days a week for many months, with few if any periods of exertion that are flat out, either in strength or speed. Each activity is, in turn, necessarily aerobic — an activity performed in balance with oxygen intake — and consequently requires that the heart can pump blood to the working muscles at several times its resting rate throughout the long duration of the exercise; also that the lungs can adequately oxygenate this enhanced blood flow as long as the exercise continues. ‘Cardio-respiratory fitness’ is thus a common feature of all endurance events, though they differ in the skeletal muscles used, and the movement patterns these muscles perform.

When muscles have been endurance-trained they are typically only a little larger than before the training began, months or years before. They become furnished, however, with a much more copious system of blood capillaries. Within the muscle fibres, mitochondria, the organelles involved in oxidative energy provision, may be 2-3 times more numerous than in untrained or differently trained fibres. Connective tissues within the muscle as well as the associated tendons and ligaments are stronger too. The nervous system must also participate in the training, for patterns of movement in the exercise concerned are usually measurably more economical than before the regime began.

Other forms of training

Pure strength training contrasts most markedly with the low-force, multiple-repetition work just described. Though increasing the bulk of the muscles and the maximum loads which they can handle, it adds little or nothing to their endurance. However the more commonly undertaken ‘weight training’, in which less extreme loads are worked against, with several times as many repetitions during the course of each gymnasium session, imparts ‘strength endurance’, a balance between the two extremes which arguably develops the most useful form of fitness for everyday life. Speed training, ‘plyometric’ (resilience) training, and flexibility training are other forms in which it is possible to specialize: in particular, yoga places a degree of emphasis upon flexibility which most other schools of physical educators would consider disproportionate. Nevertheless a programme of muscle stretching and joint flexibility should be part of the regime of every sportsperson seeking to improve not only performance but resistance to injury. Finally, between speed and endurance comes ‘anaerobic endurance’ — the ability to maintain a power output only a few per cent below flat out for several tens of seconds (as in 400 metre running) or to repeat short bursts many times in a period of about 90 min (as in hockey, soccer, and other ‘multiple sprint’ sports).

Specific versus general fitness

It would be widely agreed that the broader-based forms of fitness are of greater value in daily life than the extreme forms, such as pure endurance, pure strength, pure flexibility, or pure speed. Older literature embodied the ideal of breadth in the term ‘general fitness’. However, it is now appreciated that the dominating principle underlying the response of the body to training is its ‘specificity’. A particular exercise elicits the adaptive responses we call ‘training’ only from the specific muscles and other tissues exercised, and enhances only the specific property (endurance, strength, speed, or extensibility) which the exercise challenges. At best only very modest improvements of other properties or at other muscle sites (‘cross-training’) are ever reported, and they cannot be counted upon. A sport requiring many forms of fitness must thus have a training programme including many elements. There is probably only one sense in which ‘general fitness’ can be enhanced by most individual forms of exercise, pursued in isolation: since it is impossible to undertake any exercise without raising both pulse rate and ventilation, every form of exercise provides some cardio-respiratory training, and hence some degree of ‘general fitness’ in respect of these central organs. More thorough-going general fitness can only be attained by an exercise programme which is itself broad-based.

A broad-based programme can, of course, be achieved by regular visits to a well-conducted gymnasium; however, such a clinically purposeful regime is not the only way. Someone who, in a typical 2-week period, goes for a 40-minute run, plays a game of squash, spends an active 30 minutes in the swimming pool, does a couple of hours' heavy gardening, polishes the car energetically, chops wood, vacuum cleans the stairs twice, and scrubs the steps, especially if (s) he precedes at least the first three of these activities with 5-7 minutes of stretching and flexing exercises, will be as fit for life as a neighbour who visits the local gym three times a week. Any difference between them which is non-genetic may well be determined by which of them gets more sleep, or eats less fat.

Women, children, and the elderly

In modern, Western societies, women, children, and the elderly are particularly prone to take insufficient exercise. The Allied Dunbar National Fitness Survey found that, in England during 1990, only one woman in ten, whether aged 20 or 50, took the amount of exercise really recommended for health whereas, among the men, 30% of 20-year-olds and 20% of 50-year-olds did so. Dunbar's standards were admittedly high — among the 20-year-olds, for instance, it hoped to see three games of squash, or equivalent, per week. More recent research has shown that statistically demonstrable improvements in cardiovascular fitness, compared with the effects of taking no exercise at all, can be had from only three 20-30 minute periods per week of moderately vigorous walking. Nevertheless, about a quarter of women in the working age-groups do not even achieve this, which is a much more modest goal than the vibrant fitness sought by Dunbar.

Modern children are distracted by television and computer games and are more likely to be transported to and from school, so that they almost certainly take less exercise than their predecessors before the 1939-45 war (although incontrovertible figures for the past are hard to establish). They should be urged to the maximum amount of physical activity of which they seem capable. No damage will accrue, provided they wear well-fitting trainers, are provided with shock-absorbing landing mats for gymnastics, and don't spend more than 90 minutes, 3 days a week, with specialist, competitive coaches.

Amongst the elderly, a ‘disuse-disability spiral’ operates. Well-meaning younger carers can be the old person's worst enemies. If daily activities fail to maintain independence — the bottle top, the heavy kettle, and worst of all independence at the toilet, being critical markers of diminished capacity — exercise regimes can be of enormous benefit. Often this benefit is proportionately greater than in younger adults, because, through disuse, the elderly have declined further below their genetic capability. Instances of elderly people running marathons are well known, but strength training is at least as effective in the very old as endurance training, and may be even more beneficial.

— Neil Spurway

Bibliography

  • Morris, J. et al., (1992). Allied Dunbar National Fitness Survey. The Sports Council, London.
  • Sharkey, B. J. (1990). Physiology of fitness, (3rd edn). Human Kinetics, Champaign, Illinois.
  • Wilmore, J. H. and Costill, D. L. (2000). Physiology of sport and exercise. 2nd ed. Human Kinetics, Champaign, Illinois

See also exercise; health; sport.

Food and Fitness: fitness
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Fitness is generally defined as the ability of a person to live a happy, well-balanced life. It embraces the physical, intellectual, social, and spiritual aspects of a person's life. It is a relative term, depending on individual circumstances and for what a person needs to be fit. Fitness has health-related components and skill-related components. Health-related components include aerobic fitness, muscular strength, muscular endurance, flexibility, and body composition. Skill-related components include agility, balance, coordination, speed, power, and reaction time. Both sets of components interact and are interdependent. A deficiency in any component reduces overall fitness. Compare unfitness; see also physical fitness.

Thesaurus: fitness
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noun

  1. The quality or state of being eligible: eligibility, qualification, suitability, suitableness, worthiness. See ability/inability.
  2. A state of sound readiness: condition, fettle, form, kilter, order, shape, trim. See better/worse.

Antonyms: fitness
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n

Definition: appropriateness
Antonyms: inappropriateness, unfitness, unsuitability

n

Definition: good condition
Antonyms: poor health, unfitness


The ability to live a happy and well-balanced life. Fitness involves not only physical factors, but it also has intellectual, emotional, social, and spiritual components. These components interact and are interdependent so that if any component deviates from normal it affects the overall fitness and ability of an individual to meet the demands made by his or her way of life. Clearly, fitness is a relative term that depends on an individual's circumstances and aspirations. Fitness is also specific to a particular physical activity: a person fit to run the marathon will not necessarily be fit to do gymnastics (see physical fitness).

Good health; equine and canine sports medicine devote much enthusiasm to the measurement of fitness, the ability to perform work, or physical activity, well. Besides actual tests of performance there are biochemical, hematological and electrocardiographic tests which can be of assistance. Radiographic examination of skeletal features has special application to racing animals. Includes fitness for performing, racing or for draft work.

  • f. prediction — the art of predicting whether a horse will win a particular race; a combination of knowing the horse's racing form, fitness, owner and trainer intentions, jockey competence, suitability of the track.
 
Blogs: Related blogs on: fitness
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Wikipedia: Fitness (biology)
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Fitness (often denoted w in population genetics models) is a central idea in evolutionary theory. It describes the capability of an individual of certain genotype to reproduce, and usually is equal to the proportion of the individual's genes in all the genes of the next generation. If differences in individual genotypes affect fitness, then the frequencies of the genotypes will change over generations; the genotypes with higher fitness become more common. This process is called natural selection.

An individual's fitness is manifested through its phenotype. As phenotype is affected by both genes and environment, the fitnesses of different individuals with the same genotype are not necessarily equal, but depend on the environment in which the individuals live. However, since the fitness of the genotype is an averaged quantity, it will reflect the reproductive outcomes of all individuals with that genotype.

As fitness measures the quantity of the copies of the genes of an individual in the next generation, it doesn't really matter how the genes arrive in the next generation. That is, for an individual it is equally "beneficial" to reproduce itself, or to help relatives with similar genes to reproduce, as long as similar amount of copies of individual's genes get passed on to the next generation. Selection which promotes this kind of helper behavior is called kin selection.

The concept is particularly difficult to understand and frequently misunderstood; J.B.S. Haldane when discussing it with John Maynard Smith is reported to have described it as "a bugger".


Contents

Measures of fitness

There are two commonly used measures of fitness; absolute fitness and relative fitness.

Absolute fitness

Absolute fitness (wabs) of a genotype is defined as the ratio between the number of individuals with that genotype after selection to those before selection. It is calculated for a single generation and may be calculated from absolute numbers or from frequencies. When the fitness is larger than 1.0, the genotype increases in frequency; a ratio smaller than 1.0 indicates a decrease in frequency.

{w_{\mathrm{abs}}} = {{N_{\mathrm{after}}} \over {N_{\mathrm{before}}}}

Absolute fitness for a genotype can also be calculated as the product of the proportion survival times the average fecundity.

Relative fitness

Relative fitness is quantified as the average number of surviving progeny of a particular genotype compared with average number of surviving progeny of competing genotypes after a single generation, i.e. one genotype is normalized at w = 1 and the fitnesses of other genotypes are measured with respect to that genotype. Relative fitness can therefore take any nonnegative value, including 0.

While researchers can usually measure relative fitness, absolute fitness is more difficult. It is often difficult to determine how many individuals of a genotype there were immediately after reproduction.

The two concepts are related, and both of them are equivalent when they are divided by the mean fitness, which is weighted by genotype frequencies.

{\frac{w_{abs}}{\overline{w}_{abs}} = \frac{w_{rel}}{\overline{w}_{rel}}}

Because fitness is a coefficient, and a variable may be multiplied by it several times, biologists may work with "log fitness" (particularly so before the advent of computers). By taking the logarithm of fitness each term may be added rather than multiplied. A fitness landscape, first conceptualized by Sewall Wright, is a way of visualising fitness in terms of a three-dimensional surface on which peaks correspond to local fitness maxima; it is often said that natural selection always progresses uphill but can only do so locally. This can result in suboptimal local maxima becoming stable, because natural selection cannot return to the less-fit "valleys" of the landscape on the way to reach higher peaks.

The related concept of genetic load measures the overall fitness of a population of individuals of many genotypes whose fitnesses vary, relative to a hypothetical population in which the most fit genotype has become fixed.

Maynard-Smith's Definition

As another example we may mention the definition of fitness given by Maynard Smith in the following way: "Fitness is a property, not of an individual, but of a class of individuals – for example homozygous for allele A at a particular locus. Thus the phrase ’expected number of offspring’ means the average number, not the number produced by some one individual. If the first human infant with a gene for levitation were struck by lightning in its pram, this would not prove the new genotype to have low fitness, but only that the particular child was unlucky." [1] This measure is certainly useful in breeding programs, but hardly as a basis of a model of an evolution selecting individuals, because evolution would hardly know if the individual may be selected or not.

Hartl's Definition

Yet another possible measure has been formulated: "The fitness of the individual - having an array x of phenotypes - is the probability, s(x), that the individual will be included among the group selected as parents of the next generation." Then, the mean fitness may be determined as a mean over the set of individuals in a large population.

 P(m) = \int s(x) N(m - x)\, dx [2]

where N is the probability distribution function of phenotypes in the population, and m is its centre of gravity. This measure is a suitable basis of a model of an evolution selecting individuals. It may in principle take even the stroke of the lightning into consideration. In the case N is a Gaussian it is fairly easily proved that the average information (information entropy, disorder, diversity) of a large population may be maximized by Gaussian adaptation - keeping the mean fitness constant - in accordance with recapitulation, the central limit theorem, the Hardy-Weinberg law and the second law of thermodynamics. This is in contrast to Fisher's fundamental theorem of natural selection.

History

The British sociologist Herbert Spencer coined the phrase "survival of the fittest" (though originally, and perhaps more accurately, "survival of the best fitted") in his 1851 work Social Statics and later used it to characterise what Charles Darwin had called natural selection. The British biologist J.B.S. Haldane was the first to quantify fitness, in terms of the modern evolutionary synthesis of Darwinism and Mendelian genetics starting with his 1924 paper A Mathematical Theory of Natural and Artificial Selection. The next further advance was the introduction of the concept of inclusive fitness by the British biologist W.D. Hamilton in 1964 in his paper on The Evolution of Social Behavior.

Notes

  1. ^ Maynard-Smith, J. (1989) Evolutionary Genetics ISBN 0198542151
  2. ^ Hartl, D. L. (1981) A Primer of Population Genetics ISBN 0878932712

Further reading

  • Sober, E. (2001). The Two Faces of Fitness. In R. Singh, D. Paul, C. Krimbas, and J. Beatty (Eds.), Thinking about Evolution: Historical, Philosophical, and Political Perspectives. Cambridge University Press, pp.309-321. Full text
  • Orr HA (August 2009). "Fitness and its role in evolutionary genetics". Nat. Rev. Genet. 10 (8): 531–9. doi:10.1038/nrg2603. PMID 19546856. 

See also

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